Vegetative features similar to Equisetales; Reproductive parts are similar to the Sphenophyllales
Distinct nodes and internodes
Basal nodes produce 1-2 first-order branches up to 3 m long
Second-order branches are opposite and decussate
Higher-order branches emerge in pseudo-distichous pattern
Pseudomonopodial or monopodial branching
15-20 meters tall
Anatomy is unknown
Originally called meiophylls
Produced only on high-order branches
Highly dissected leaflet attached to twice-dichotomous petiole
Underground portions unknown
Fertile branches up to 30 cm long
Each fertile unit exhibits whorled bracts and sporangiophores
Sporangiophores have upturned tips divided into two segments
Each sporangiophore has ~30 sporangia
└Equisetophytes
└Pseudoborniales †
Late Devonian
Pseudobornia ursina † (Schweitzer 1967)